Bearded Collie


    The Bearded Collie, or Beardie, is a herding breed of dog once used primarily by Scottish shepherds, but now mostly a popular family companion.
    Bearded Collies have an average weight of 18–27 kilograms (40–60 lb). Males are around 53–56 centimetres (21–22 in) tall at the withers while females are around 51–53 centimetres (20–21 in) tall

History:
    The Bearded Collie's history is a combination of fact and legend. Kazimierz Grabski, a Polish merchant, reportedly traded a shipment of grain for sheep in Scotland in 1514 and brought six Polish Lowland Sheepdogs to move them. A Scottish shepherd was so impressed with the herding ability of the dogs that he traded several sheep for several dogs. The Polish sheepdogs were bred with local Scottish dogs to produce the Bearded Collie.
It is generally agreed that Mrs. G. Olive Willison founded the modern Bearded Collie in 1944 with her brown bitch, Jeannie of Bothkennar. Jeannie was supposedly a Shetland Sheepdog, but Mrs. Willison received a Bearded Collie by accident. She was so fascinated by the dog that she wanted to begin breeding, so she began searching for a dog for Jeannie. While walking along the beach, Mrs. Willison met a man who was emigrating from Scotland; she became the owner of his grey dog, David, who became Bailie of Bothkennar.
Bailie and Jeannie of Bothkennar are the founders of the modern breed; there are only a few other registrable blood lines, preserved in large part by the perseverance of Mr. Nicolas Broadbridge (Sallen) and Mrs. Betty Foster (Bredon). These are based on Turnbull's Blue—a Bearded Collie from pure working stock, registered in ISDS when ISDS still registered non-Border Collies. He sired three litters of registerable Bearded Collies.

Working life:
    The Bearded Collie is used to herd both sheep and cattle. It is essentially a working dog—bred to be hardy and reliable, able to stand up to the harshest conditions and the toughest sheep. The working Bearded Collie has become less common in the last few decades and might have died out; however, thanks to the efforts of a few shepherds like Tom Muirhead and Peter Wood, the "working Beardie" has survived and is becoming more popular. It has been exported to Australia and the United States, and finds favour among those looking for an independent and intelligent sheepdog. The Working Bearded Collie Society's mission is to preserve the working abilities of non-registered working dogs from "bearded" ancestors. The website Shepherds with beardies has much valuable information on the small population of working Beardies.
    The KC-registered Bearded Collie has fallen into disfavour with the shepherds of Wales, Scotland and elsewhere because of the show-breeding community's lack of interest in producing "hardy and reliable" animals; show-bred lines tend to develop excessive coats, in particular. However, in some countries (notably Sweden and the United States) herding programmes have been developed for the breed. The breed organisations in those countries actively encourage breeders to emphasise qualities other than appearance.
    The Bearded Collie may have earned its nickname "bouncing Beardie" because the dogs would work in thick underbrush on hillsides; they would bounce to catch sight of the sheep. Beardies also have a characteristic way of facing a stubborn ewe, barking and bouncing on the forelegs. Whatever the reason, a typical Bearded Collie is an enthusiastic herding dog which requires structure and care; it moves stock with body, bark and bounce as required. Very few Beardies show "eye" when working; most are upright.

Herding instincts and tractability can be assessed in noncompetitive herding tests. Beardies exhibiting basic herding instincts can be trained to compete in herding trials.
    The breed became popular during the last half of the 20th century—propelled, in part, by Potterdale Classic at Moonhill, a Bearded Collie who won Best in Show at Crufts in 1989. The Bearded Collie Club celebrated its Golden Jubilee in 2005.The bearded collie is also very good natured and is good as a family pet and a working dog and a show dog.



Bear


    Bears are mammals of the family Ursidae. Bears are classified as caniforms, or doglike carnivorans, with the pinnipeds being their closest living relatives. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats throughout the Northern Hemisphere and partially in the Southern Hemisphere. Bears are found on the continents of North America, South America, Europe, and Asia.
    Common characteristics of modern bears include large bodies with stocky legs, long snouts, shaggy hair, plantigrade paws with five nonretractile claws, and short tails. While the polar bear is mostly carnivorous, and the giant panda feeds almost entirely on bamboo, the remaining six species are omnivorous with varied diets.
    With the exception of courting individuals and mothers with their young, bears are typically solitary animals. They are generally diurnal, but may be active during the night (nocturnal) or twilight (crepuscular), particularly around humans. Bears possess an excellent sense of smell and, despite their heavy build and awkward gait, are adept runners, climbers, and swimmers. Bears use shelters, such as caves and burrows, as their dens; most species occupy their dens during the winter for a long period (up to 100 days) of sleep similar to hibernation.
    Bears have been hunted since prehistoric times for their meat and fur. With their tremendous physical presence and charisma, they play a prominent role in the arts, mythology, and other cultural aspects of various human societies. In modern times, the bears' existence has been pressured through the encroachment on their habitats and the illegal trade of bears and bear parts, including the Asian bile bear market. The IUCN lists six bear species as vulnerable or endangered, and even least concern species, such as the brown bear, are at risk of extirpation in certain countries. The poaching and international trade of these most threatened populations are prohibited, but still ongoing.

Morphology:
    Bears are generally bulky and robust animals with relatively short legs. They are sexually dimorphic with regard to size, with the males being larger. Larger species tend to show increased levels of sexual dimorphism in comparison to smaller species and where a species varies in size across its distribution, individuals from larger-sized areas tend also to vary more. Bears include the most massive terrestrial members of the order Carnivora. Some exceptional polar bears and Kodiak bears (a brown bear subspecies) have been weighed at 1,002 kg (2,209 lb) and 751 kg (1,656 lb) respectively. As to which species is the largest depends on whether the assessment is based on which species has the largest individuals (brown bears) or on the largest average size (polar bears), as some races of brown bears are much smaller than polar bears. Adult male Kodiak bears average 480 to 533 kg (1,058 to 1,175 lb) compared to an average of 386 to 408 kg (851 to 899 lb) in adult male polar bears, per the Guinness Book of World Records. The smallest bears are the sun bears of Asia, which weigh an average of 65 kg (143 lb) for the males and 45 kg (99 lb) for the females, though the smallest mature females can weigh only 20 kg (44 lb). All "medium"-sized bear species (which include the other five extant species) are around the same average weight, with males averaging around 100 to 120 kg (220 to 260 lb) and females averaging around 60 to 85 kg (132 to 187 lb), although it is not uncommon for male American black bears to considerably exceed "average" weights. Head-and-body length can range from 120 cm (47 in) in sun bears to 300 cm (120 in) in large polar and brown bears and shoulder height can range from 60 cm (24 in) to over 160 cm (63 in) in the same species, respectively. The tails of bears are often considered a vestigial feature and can range from 3 to 22 cm (1.2 to 8.7 in).

Unlike most other land carnivorans, bears are plantigrade. They distribute their weight toward the hind feet, which makes them look lumbering when they walk. They are still quite fast, with the brown bear reaching 48 km/h (30 mph), although they are still slower than felines and canines. Bears can stand on their hind feet and sit up straight with remarkable balance. Bears' nonretractable claws are used for digging, climbing, tearing, and catching prey. Their ears are rounded.
    Bears have an excellent sense of smell, better than the dogs (Canidae), or possibly any other mammal. This sense of smell is used for signalling between bears (either to warn off rivals or detect mates) and for finding food. Smell is the principal sense used by bears to find most of their food.

Distribution and habitat:
    Bears are primarily found in the Northern Hemisphere, and with one exception, only in Asia, North America, and Europe. The single exception is the spectacled bear (Tremarctos ornatus); native to South America it inhabits the Andean region. The Atlas bear, a subspecies of the brown bear, was the only bear native to Africa. It was distributed in North Africa from Morocco to Libya, but has been extinct since around the 1870s. The most widespread species is the brown bear, which occurs from Western Europe eastwards through Asia to the western areas of North America. The American black bear is restricted to North America, and the polar bear is restricted to the Arctic Sea. All the remaining species are Asian.
    With the exception of the polar bear, bears are mostly forest species. Some species, particularly the brown bear, may inhabit or seasonally use other areas, such as alpine scrub or tundra.

Behaviour:
    Bears are generally diurnal, meaning that they are active for the most part during the day. The belief that they are nocturnal apparently comes from the habits of bears that live near humans, which engage in some nocturnal activities, such as raiding trash cans or crops while avoiding humans. The sloth bear of Asia is the most nocturnal of the bears, but this varies by individual, and females with cubs are often diurnal to avoid competition with males and nocturnal predators. Bears are overwhelmingly solitary and are considered to be the most asocial of all the Carnivora. Liaisons between breeding bears are brief, and the only times bears are encountered in small groups are mothers with young or occasional seasonal bounties of rich food (such as salmon runs


Beagle


    The Beagle is a breed of small-sized hound, similar in appearance to the much larger foxhound. The Beagle is a scent hound, developed primarily for hunting hare. With a great sense of smell and their tracking instinct, the Beagle is employed as detection dog for prohibited agricultural imports and foodstuffs in quarantine around the world. The Beagle is intelligent but single-minded. It is a popular pet due to its size, good temper, and lack of inherited health problems.
    Although beagle-type dogs have existed for 2,500 years, the modern breed was developed in Great Britain around the 1830s from several breeds, including the Talbot Hound, the North Country Beagle, the Southern Hound, and possibly the Harrier.
    Beagles have been depicted in popular culture since Elizabethan times in literature and paintings, and more recently in film, television, and comic books. Snoopy of the comic strip Peanuts has been promoted as "the world's most famous beagle".

History
Dogs of similar size and purpose to the modern Beagle[a] can be traced in Ancient Greece[2] back to around the 5th century BC. Xenophon, born around 430 BC, in his Treatise on Hunting or Cynegeticus refers to a hound that hunted hares by scent and was followed on foot. Small hounds are mentioned in the Forest Laws of Canute which exempted them from the ordinance which commanded that all dogs capable of running down a stag should have one foot mutilated.[3] If genuine, these laws would confirm that beagle-type dogs were present in England before 1016, but it is likely the laws were written in the Middle Ages to give a sense of antiquity and tradition to Forest Law.[4]


The Southern Hound is thought to be an ancestor of the Beagle
In the 11th century, William the Conqueror brought the Talbot hound to Britain. The Talbot was a predominantly white, slow, deep-throated, scent hound derived from the St. Hubert Hound which had been developed in the 8th century. At some point the English Talbots were crossed with Greyhounds to give them an extra turn of speed.[5] Long extinct, the Talbot strain probably gave rise to the Southern Hound which, in turn, is thought to be an ancestor of the modern-day Beagle.[b]

From medieval times, beagle was used as a generic description for the smaller hounds, though these dogs differed considerably from the modern breed. Miniature breeds of beagle-type dogs were known from the times of Edward II and Henry VII, who both had packs of Glove Beagles, so named since they were small enough to fit on a glove, and Queen Elizabeth I kept a breed known as a Pocket Beagle, which stood 8 to 9 inches (20 to 23 cm) at the shoulder. Small enough to fit in a "pocket" or saddlebag, they rode along on the hunt. The larger hounds would run the prey to ground, then the hunters would release the small dogs to continue the chase through underbrush. Elizabeth I referred to the dogs as her singing beagles and often entertained guests at her royal table by letting her Pocket Beagles cavort amid their plates and cups.[6] 19th-century sources refer to these breeds interchangeably and it is possible that the two names refer to the same small variety. In George Jesse's Researches into the History of the British Dog from 1866, the early 17th-century poet and writer Gervase Markham is quoted referring to the Beagle as small enough to sit on a man's hand and to the:

little small mitten-beagle, which may be companion for a ladies kirtle, and in the field will run as cunningly as any hound whatere, only their musick is very small like reeds.[7]

Standards for the Pocket Beagle were drawn up as late as 1901; these genetic lines are now extinct, although modern breeders have attempted to recreate the variety.[8]


This image from the turn of the 19th century shows a dog with a heavier body and lacking the features of later strains.
By the 18th century two breeds had been developed for hunting hare and rabbit: the Southern Hound and the North Country Beagle (or Northern Hound). The Southern Hound, a tall, heavy dog with a square head, and long, soft ears, was common from south of the River Trent and probably closely related to the Talbot Hound. Though slow, it had stamina and an excellent scenting ability. The North Country Beagle, possibly a cross between an offshoot of the Talbot stock and a Greyhound, was bred chiefly in Yorkshire and was common in the northern counties. It was smaller than the Southern Hound, less heavy-set and with a more pointed muzzle. It was faster than its southern counterpart but its scenting abilities were less well developed.[9] As fox hunting became increasingly popular, numbers of both types of hound diminished. The beagle-type dogs were crossed with larger breeds such as Stag Hounds to produce the modern Foxhound. The beagle-size varieties came close to extinction but some farmers in the South ensured the survival of the prototype breeds by maintaining small rabbit-hunting packs.

Development of the modern breed
Reverend Phillip Honeywood established a Beagle pack in Essex in the 1830s and it is believed that this pack formed the basis for the modern Beagle breed. Although details of the pack's lineage are not recorded it is thought that North Country Beagles and Southern Hounds were strongly represented; William Youatt suspected that Harriers formed a good majority of the Beagle's bloodline, but the origin of the Harrier is itself obscure.[10] Honeywood's Beagles were small, standing at about 10 inches (25 cm) at the shoulder, and pure white according to John Mills (writing in The Sportsman's Library in 1845). Prince Albert and Lord Winterton also had Beagle packs around this time, and royal favour no doubt led to some revival of interest in the breed, but Honeywood's pack was regarded as the finest of the three.[11]


Early images of the Beagle (clockwise from top left): 1833, 1835, Stonehenge's Medium (1859, reusing Youatt's 1852 "Beagle" image) and Dwarf Beagle (1859).
Although credited with the development of the modern breed, Honeywood concentrated on producing dogs for hunting and it was left to Thomas Johnson to refine the breeding to produce dogs that were both attractive and capable hunters. Two strains were developed: the rough- and smooth-coated varieties. The rough-coated Beagle survived until the beginning of the 20th century, and there were even records of one making an appearance at a dog show as late as 1969, but this variety is now extinct, having probably been absorbed into the standard Beagle bloodline.[12]

In the 1840s, a standard Beagle type was beginning to develop; the distinction between the North Country Beagle and Southern Hound had been lost, but there was still a large variation in size, character, and reliability among the emerging packs.[13] In 1856, "Stonehenge" (the pseudonym of John Henry Walsh), writing in the Manual of British Rural Sports, was still dividing Beagles into four varieties: the medium Beagle; the dwarf or lapdog Beagle; the fox Beagle (a smaller, slower version of the Foxhound); and the rough-coated or terrier Beagle, which he classified as a cross between any of the other varieties and one of the Scottish terrier breeds.[14] Stonehenge also gives the start of a standard description:

In size the beagle measures from 10 inches, or even less, to 15. In shape they resemble the old southern hound in miniature, but with more neatness and beauty; and they also resemble that hound in style of hunting.[14]

By 1887 the threat of extinction was on the wane: there were 18 Beagle packs in England.[15] The Beagle Club was formed in 1890 and the first standard drawn up at the same time.[16] The following year the Association of Masters of Harriers and Beagles was formed. Both organisations aimed to further the best interests of the breed, and both were keen to produce a standard type of Beagle.[17] By 1902, the number of packs had risen to 44.[15]

Export
Beagles were in the United States by the 1840s at the latest, but the first dogs were imported strictly for hunting and were of variable quality. Since Honeywood had only started breeding in the 1830s, it is unlikely these dogs were representative of the modern breed and the description of them as looking like straight-legged Dachshunds with weak heads has little resemblance to the standard. Serious attempts at establishing a quality bloodline began in the early 1870s when General Richard Rowett from Illinois imported some dogs from England and began breeding. Rowett's Beagles are believed to have formed the models for the first American standard, drawn up by Rowett, L. H. Twadell, and Norman Ellmore in 1887.[18] The Beagle was accepted as a breed by the American Kennel Club (AKC) in 1885.[19] In the 20th century the breed has spread worldwide.

Popularity

An attractive uniform type for the breed developed at the start of the 20th century
On its formation, the Association of Masters of Harriers and Beagles took over the running of a regular show at Peterborough that had started in 1889, and the Beagle Club in the UK held its first show in 1896.[16] The regular showing of the breed led to the development of a uniform type, and the Beagle continued to prove a success up until the outbreak of World War I when all shows were suspended. After the war, the breed was again struggling for survival in the UK: the last of the Pocket Beagles was probably lost during this time, and registrations fell to an all-time low. A few breeders (notably Reynalton Kennels) managed to revive interest in the dog and by World War II, the breed was once again doing well. Registrations dropped again after the end of the war but almost immediately recovered.[20]

As purebred dogs, Beagles have always been more popular in the United States and Canada than in their native country England. The National Beagle Club of America was formed in 1888 and by 1901 a Beagle had won a Best in Show title. As in the UK, activity during World War I was minimal, but the breed showed a much stronger revival in the U.S. when hostilities ceased. In 1928 it won a number of prizes at the Westminster Kennel Club's show and by 1939 a Beagle – Champion Meadowlark Draughtsman – had captured the title of top-winning American-bred dog for the year.[21] On 12 February 2008, a Beagle, K-Run's Park Me In First (Uno), won the Best In Show category at the Westminster Kennel Club show for the first time in the competition's history.[22] In North America they have been consistently in the top-ten most-popular breeds for over 30 years. From 1953 to 1959 the Beagle was ranked No. 1 on the list of the American Kennel Club's registered breeds;[23] in 2005 and 2006 it ranked 5th out of the 155 breeds registered.[24] In the UK they are not quite so popular, placing 28th and 30th in the rankings of registrations with the Kennel Club in 2005 and 2006 respectively.[25] In the United States the Beagle ranked 4th most popular breed in 2012 and 2013, behind the Labrador Retriever (#1), German Shepherd (#2) and Golden Retriever (#3) breeds.[26]

Name
According to the Oxford English Dictionary, the first mention of the beagle by name in English literature dates from c. 1475 in The Squire of Low Degree. The origin of the word "beagle" is uncertain, although it has been suggested that the word derives from the French begueule.[27] (meaning "wide throat" from bayer "open wide" and gueule "mouth")

It is not known why the black and tan Kerry Beagle, present in Ireland since Celtic times, has the beagle description, since at 22 to 24 inches (56 to 61 cm) it is significantly taller than the modern day Beagle, and in earlier times was even larger. Some writers suggest that the Beagle's scenting ability may have come from cross-breeding earlier strains with the Kerry Beagle. Originally used for hunting stags, it is today used for hare and drag hunting.[28]

Appearance

The Kennel Club (UK) standard states the Beagle should give the impression of quality without coarseness.
The general appearance of the Beagle resembles a miniature Foxhound, but the head is broader and the muzzle shorter, the expression completely different and the legs shorter in proportion to the body.[29] They are generally between 13 and 16 inches (33 and 41 cm) high at the withers and weigh between 18 and 35 lb (8.2 and 15.9 kg), with females being slightly smaller than males on average.[30]

They have a smooth, somewhat domed skull with a medium-length, square-cut muzzle and a black (or occasionally liver) gumdrop nose. The jaw is strong and the teeth scissor together with the upper teeth fitting perfectly over the lower teeth and both sets aligned square to the jaw. The eyes are large, hazel or brown, with a mild hound-like pleading look. The large ears are long, soft and low-set, turning towards the cheeks slightly and rounded at the tips. Beagles have a strong, medium-length neck (which is long enough for them to easily bend to the ground to pick up a scent), with little folding in the skin but some evidence of a dewlap; a broad chest narrowing to a tapered abdomen and waist and a long, slightly curved tail (known as the "stern") tipped with white. The white tip, known as the flag has been selectively bred for, as it allows the dog to be easily seen when its head is down following a scent.[31] The tail does not curl over the back, but is held upright when the dog is active. The Beagle has a muscular body and a medium-length, smooth, hard coat. The front legs are straight and carried under the body while the rear legs are muscular and well bent at the stifles.[32]

The tricolored Beagle—white with large black areas and light brown shading—is the most common. Tricolored Beagles occur in a number of shades, from the "Classic Tri" with a jet black saddle (also known as "Blackback"), to the "Dark Tri" (where faint brown markings are intermingled with more prominent black markings), to the "Faded Tri" (where faint black markings are intermingled with more prominent brown markings). Some tricolored dogs have a broken pattern, sometimes referred to as pied. These dogs have mostly white coats with patches of black and brown hair. Tricolor Beagles are almost always born black and white. The white areas are typically set by eight weeks, but the black areas may fade to brown as the puppy matures. (The brown may take between one and two years to fully develop.) Some Beagles gradually change color during their lives, and may lose their black markings entirely.

Two-color varieties always have a white base color with areas of the second color. Tan and white is the most common two-color variety, but there is a wide range of other colors including lemon, a very light tan; red, a reddish, almost orange, brown; and liver, a darker brown, and black. Liver is not common and is not permitted in some standards; it tends to occur with yellow eyes. Ticked or mottled varieties may be either white or black with different colored flecks (ticking), such as the blue-mottled or bluetick Beagle, which has spots that appear to be a midnight-blue color, similar to the coloring of the Bluetick Coonhound. Some tricolor Beagles also have ticking of various colors in their white areas.[33][34]

Sense of smell
Alongside the Bloodhound and Basset Hound, the Beagle has one of the best developed senses of smell of any dog.[35] In the 1950s, John Paul Scott and John Fuller began a 13-year study of canine behavior. As part of this research, they tested the scenting abilities of various breeds by putting a mouse in a one-acre field and timing how long it took the dogs to find it. The Beagles found it in less than a minute, while Fox Terriers took 15 minutes and Scottish Terriers failed to find it at all. Beagles are better at ground-scenting (following a trail on the ground) than they are at air-scenting, and for this reason they have been excluded from most mountain rescue teams in favor of collies, which use sight in addition to air-scenting and are more biddable.[35] The long ears and large lips of the Beagle probably assist in trapping the scents close to the nose.[36]

Variations
Breed varieties
The American Kennel Club recognizes two separate varieties of Beagle: the 13-inch for hounds less than 13 inches (33 cm), and the 15-inch for those between 13 and 15 inches (33 and 38 cm). The Canadian Kennel Club recognizes a single type, with a height not exceeding 15 inches (38 cm). The Kennel Club (UK) and FCI affiliated clubs recognize a single type, with a height of between 13 and 16 inches (33 and 41 cm).


A Puggle, a Beagle/Pug cross, shows traits from both breeds.
English and American varieties are sometimes mentioned. However, there is no official recognition from any Kennel Club for this distinction. Beagles fitting the American Kennel Club standard – which disallows animals over 15 inches (38 cm) – are smaller on average than those fitting the Kennel Club standard which allows heights up to 16 inches (41 cm).

Pocket Beagles are sometimes advertised for sale but while the UK Kennel Club originally specified a standard for the Pocket Beagle in 1901, the variety is not now recognized by any Kennel Club. A strain known as Patch Hounds was developed by Willet Randall and his family from 1896 specifically for their rabbit hunting ability. They trace their bloodline back to Field Champion Patch, but do not necessarily have a patchwork marking.[37][38]

Crossbreeds
In the 1850s, Stonehenge recommended a cross between a Beagle and a Scottish Terrier as a retriever. He found the crossbreed to be a good worker, silent and obedient, but it had the drawback that it was small and could barely carry a hare.[39]

More recently the trend has been for "designer dogs" and one of the most popular has been the Beagle/Pug cross known as a Puggle. Some puppies of this cross are less excitable than a Beagle and with a lower exercise requirement, similar to the Pug parent; but many are highly excitable and require vigorous exercise.[40]

Temperament

Beagles are happy to rest without being exercised to exhaustion.
The Beagle has an even temper and gentle disposition. Described in several breed standards as "merry", they are amiable and typically neither aggressive nor timid, although this depends on the individual. They enjoy company, and although they may initially be standoffish with strangers, they are easily won over. They make poor guard dogs for this reason, although their tendency to bark or howl when confronted with the unfamiliar makes them good watch dogs. In a 1985 study conducted by Ben and Lynette Hart, the Beagle was given the highest excitability rating, along with the Yorkshire Terrier, Cairn Terrier, Miniature Schnauzer, West Highland White Terrier, and Fox Terrier.[41][c]

Beagles are intelligent but, as a result of being bred for the long chase, are single-minded and determined, which can make them hard to train. They can be difficult to recall once they have picked up a scent, and are easily distracted by smells around them. They do not generally feature in obedience trials; while they are alert, respond well to food-reward training, and are eager to please, they are easily bored or distracted. They are ranked 72nd in Stanley Coren's The Intelligence of Dogs, as Coren places them among the group with the lowest degree of working/obedience intelligence. Coren's scale, however, does not assess understanding, independence, or creativity.[42][43]

Beagles are excellent with children and this is one of the reasons they have become popular family pets, but they are pack animals, and are prone to separation anxiety,[44] a condition which causes them to destroy things when left unattended. Not all Beagles will howl, but most will bark when confronted with strange situations, and some will bay (also referred to as "speaking", "giving tongue", or "opening") when they catch the scent of potential quarry.[45] They also generally get along well with other dogs. They are not too demanding with regard to exercise; their inbred stamina means they do not easily tire when exercised, but they also do not need to be worked to exhaustion before they will rest. Regular exercise helps ward off the weight gain to which the breed is prone.[46]

Health

Day-old Beagle puppies
The typical longevity of Beagles is 12–15 years,[47] which is a common lifespan for dogs of their size.[48]

Beagles may be prone to epilepsy, but this can often be controlled with medication. Hypothyroidism and a number of types of dwarfism occur in Beagles. Two conditions in particular are unique to the breed: "Funny Puppy", in which the puppy is slow to develop and eventually develops weak legs, a crooked back and although normally healthy, is prone to a range of illnesses;[49] Hip dysplasia, common in Harriers and in some larger breeds, is rarely considered a problem in Beagles.[50] Beagles are considered a chondrodystrophic breed, meaning that they are prone to types of disk diseases.[51]


Weight gain can be a problem in older or sedentary dogs, which in turn can lead to heart and joint problems.
In rare cases, Beagles may develop immune mediated polygenic arthritis (where the immune system attacks the joints) even at a young age. The symptoms can sometimes be relieved by steroid treatments.[49] Another rare disease in the breed is neonatal cerebellar cortical degeneration. Affected puppies are slow, have lower co-ordination, fall more often and don't have a normal gait. It has an estimated carrier rate of 5% and affected rate of 0.1%. A genetic test is available.[52][53]

Their long floppy ears can mean that the inner ear does not receive a substantial air flow or that moist air becomes trapped, and this can lead to ear infections. Beagles may also be affected by a range of eye problems; two common ophthalmic conditions in Beagles are glaucoma and corneal dystrophy.[54] "Cherry eye", a prolapse of the gland of the third eyelid, and distichiasis, a condition in which eyelashes grow into the eye causing irritation, sometimes exist; both these conditions can be corrected with surgery.[49] They can suffer from several types of retinal atrophy. Failure of the nasolacrimal drainage system can cause dry eye or leakage of tears onto the face.[49]

As field dogs they are prone to minor injuries such as cuts and sprains, and, if inactive, obesity is a common problem as they will eat whenever food is available and rely on their owners to regulate their weight.[49] When working or running free they are also likely to pick up parasites such as fleas, ticks, harvest mites, and tapeworms, and irritants such as grass seeds can become trapped in their eyes, soft ears, or paws.[55]

Beagles may exhibit a behaviour known as reverse sneezing, in which they sound as if they are choking or gasping for breath, but are actually drawing air in through the mouth and nose. The exact cause of this behaviour is not known, but it is not harmful to the dog.[56]

Hunting
Main article: Beagling

The Caynsham Foot Beagles (c. 1885)
Beagles were developed primarily for hunting hare, an activity known as beagling. They were seen as ideal hunting companions for the elderly who could follow on horseback without exerting themselves, for young hunters who could keep up with them on ponies, and for the poorer hunters who could not afford to maintain a stable of good hunting horses.[57] Before the advent of the fashion for foxhunting in the 19th century, hunting was an all day event where the enjoyment was derived from the chase rather than the kill. In this setting the tiny Beagle was well matched to the hare, as unlike Harriers they would not quickly finish the hunt, but because of their excellent scent-tracking skills and stamina they were almost guaranteed to eventually catch the hare. The Beagle packs would run closely together ("so close that they might be covered with a sheet"[10]) which was useful in a long hunt, as it prevented stray dogs from obscuring the trail. In thick undergrowth they were also preferred to spaniels when hunting pheasant.[58]

With the fashion for faster hunts, the Beagle fell out of favour for chasing hare, but was still employed for rabbit hunting. In Anecdotes of Dogs (1846), Edward Jesse says:

In rabbit-shooting, in gorse and thick cover, nothing can be more cheerful than the beagle. They also are easily heard over long distances and in thick cover. They have been called rabbit-beagles from this employment, for which they are peculiarly qualified, especially those dogs which are somewhat wire-haired.[6]


The Beagle has been used for rabbit-hunting since the earliest development of the breed.
In the United States they appear to have been employed chiefly for hunting rabbits from the earliest imports. Hunting hare with Beagles became popular again in Britain in the mid-19th century and continued until it was made illegal in Scotland by the Protection of Wild Mammals (Scotland) Act 2002 and in England and Wales by the Hunting Act 2004. Under this legislation Beagles may still pursue rabbits with the landowner's permission. Drag hunting is popular where hunting is no longer permitted or for those owners who do not wish to participate in hunting a live animal, but still wish to exercise their dog's innate skills.

The traditional foot pack consists of up to 40 Beagles, marshaled by a Huntsman who directs the pack and who is assisted by a variable number of whippers-in whose job is to return straying hounds to the pack. The Master of the Hunt is in overall day-to-day charge of the pack, and may or may not take on the role of Huntsman on the day of the hunt.

As hunting with Beagles was seen as ideal for young people, many of the British public schools traditionally maintained Beagle packs. Protests were lodged against Eton's use of Beagles for hunting as early as 1902 but the pack is still in existence today,[59] and a pack used by Imperial College in Wye, Kent was stolen by the Animal Liberation Front in 2001.[60] School and university packs are still maintained by Eton, Marlborough, Wye, Radley, the Royal Agricultural University and Christ Church, Oxford.[61]

In addition to organized beagling, beagles have been used for hunting or flushing to guns (often in pairs) a wide range of game including snowshoe hare, cottontail rabbits, game birds, roe deer, red deer, bobcat, coyote, wild boar and foxes, and have even been recorded as being used to hunt stoat.[62][63] In most of these cases, the Beagle is employed as a gun dog, flushing game for hunter's guns.[62]

Detection

Beagles have excellent noses; this dog is employed by the US Customs and Border Protection Agency.
Beagles are used as detection dogs in the Beagle Brigade of the United States Department of Agriculture. These dogs are used to detect food items in luggage being taken into the United States. After trialling several breeds, Beagles were chosen because they are relatively small and unintimidating for people who are uncomfortable around dogs, easy to care for, intelligent and work well for rewards.[64] They are also used for this purpose in a number of other countries including by the Ministry of Agriculture and Forestry in New Zealand, the Australian Quarantine and Inspection Service, and in Canada, Japan and the People's Republic of China.[65] Larger breeds are generally used for detection of explosives as this often involves climbing over luggage and on large conveyor belts, work for which the smaller Beagle is not suited.[66]

Testing
Beagles are the dog breed most often used in animal testing, due to their size and passive nature. Beagles are used in a range of research procedures: fundamental biological research, applied human medicine, applied veterinary medicine, and protection of man, animals or the environment.[67][68] Of the 8,018 dogs used in testing in the UK in 2004, 7,799 were Beagles (97.3%).[69] In the UK, the Animals (Scientific Procedures) Act 1986 gave special status to primates, equids, cats and dogs and in 2005 the Animal Procedures Committee (set up by the act) ruled that testing on mice was preferable, even though a greater number of individual animals were involved.[70] In 2005 Beagles were involved in less than 0.3% of the total experiments on animals in the UK, but of the 7670 experiments performed on dogs 7406 involved Beagles (96.6%).[67] Most dogs are bred specifically for this purpose, by companies such as Harlan. In the UK companies breeding animals for research must be licensed under the Animals (Scientific Procedures) Act.[70]

Testing of cosmetic products on animals is banned in the member states of European Community,[71] although France protested the ban and has made efforts to have it lifted.[72] It is permitted in the United States but is not mandatory if safety can be ascertained by other methods, and the test species is not specified by the Food and Drug Administration (FDA).[73] When testing toxicity of food additives, food contaminants, and some drugs and chemicals the FDA uses Beagles and miniature pigs as surrogates for direct human testing.[74] Minnesota was the first state to enact a Beagle freedom adoption law in 2014, mandating that dogs and cats are allowed to be adopted once they have completed with research testing.[75]

Anti-vivisection groups have reported on abuse of animals inside testing facilities. In 1997 footage secretly filmed by a freelance journalist inside Huntingdon Life Sciences in the UK showed staff punching and screaming at Beagles.[76] Consort Kennels, a UK-based breeder of Beagles for testing, closed down in 1997 after pressure from animal rights groups.[77]

Other roles
Although bred for hunting, Beagles are versatile and are nowadays employed for various other roles in detection, therapy, and as family pets.[31] Beagles are used as sniffer dogs for termite detection in Australia,[78] and have been mentioned as possible candidates for drug and explosive detection.[79][80] Because of their gentle nature and unimposing build, they are also frequently used in pet therapy, visiting the sick and elderly in hospital.[81] In June 2006, a trained Beagle assistance dog was credited with saving the life of its owner after using her owner's mobile phone to dial an emergency number.[82] In the aftermath of the 2010 Haiti earthquake, a Beagle search and rescue dog with a Colombian rescue squad was credited with locating the owner of the Hôtel Montana, who was subsequently rescued after spending 100 hours buried in the rubble.[83] Beagles were hired by New York City to help with bedbug detection,[84] while the role of such dogs in this type of detection may have doubts.[85]

In popular culture
Beagles have been featured across a wide range of media. References to the dog appear before the 19th century in works by such writers as William Shakespeare, John Webster, John Dryden, Thomas Tickell, Henry Fielding, and William Cowper, as well as in Alexander Pope's translation of Homer's Iliad.[d] Beagles appeared in funny animal comic strips and animated cartoons from the 1950s with the Peanuts character Snoopy billed as "the world's most famous Beagle";[1] Walt Disney's Beagle Boys; Garfield's friend and "chew dog" Odie; and Beegle Beagle, the constant companion of Hanna-Barbera's Grape Ape and Cartoon Network's Courage the Cowardly Dog. They have appeared in numerous films, taking a central role in Underdog, Cats & Dogs and its sequel, and the title role in the adaptation of Phyllis Reynolds Naylor's book Shiloh. They have played supporting roles in films including Audition, The Monster Squad, I Am Number Four, and The Royal Tenenbaums, and on television in Star Trek: Enterprise, EastEnders, The Wonder Years, and To the Manor Born, among others.

Former US President Lyndon Baines Johnson had several Beagles, and caused an outcry when he picked up one of them by its ears during an official greeting on the White House lawn.[86] The ship on which Charles Darwin made the voyage which provided much of the inspiration for On the Origin of Species was named HMS Beagle after the breed, and, in turn, lent its name to the ill-fated British Martian lander Beagle 2. A Canadian bred 15 inch female Beagle with the registered name of Gr Ch Tashtins Lookin For Trouble and the pet name of "Miss P" won the 2015 Westminster Kennel Club Dog Show.[87]


Bavarian Mountain Hound


    The Bavarian Mountain Hound (German = "Bayerischer Gebirgsschweißhund") is a breed of dog from Germany. As a scent hound, it has been used in Germany since the early 20th century to trail wounded game. It is a cross between the Bavarian Hound and the Hanover Hound.

Appearance:
    The Bavarian Mountain Hound's head is strong and elongated. The skull is relatively broad and slightly domed. It has a pronounced stop and a slightly curved nosebridge. The muzzle should be broad with solid jaws, and its lips fully covering the mouth. Its nose is black or dark red with wide nostrils. Its ears are high set and medium in length. They are broader at the base and rounded at the tips, hanging heavily against the head. Its body is slightly longer than it is tall and slightly raised at the rump. The neck medium in length, strong, with a slight dewlap. Topline sloping slightly upward from withers to hindquarters. Chest well-developed, long, moderately wide, and well let-down with a slight tuck-up. It has a long, fairly straight croup and solid back. While its tail is set on high, medium in length and hanging to the hock, carried level to the ground or hanging down.

Size:
    Bavarian Mountain Hounds weigh between 20 and 25 kg, males are 47 to 52 cm (18.5 to 20.5 in) high, while females are 44 to 48 cm (17.5 to 19 in).

Coat and color:
    The coat is short, thick and shiny, lying very flat against the body, and moderately harsh. It is finer on the head and ears, harsher and longer on the abdomen, legs, and tail. Its coat can come in all shades of black-masked fawn or brindle.

Temperament:
    Bavarian Mountain Hounds are calm, quiet, poised, and very attached to their masters and family. When hunting, they are hard, single-minded, and persistent, courageous, spirited, fast, and agile, they are at ease on a rugged terrain, with a superb nose and powerful hunting instinct. However, they need a patient, experienced trainer.

Care:
    The Bavarian Mountain is not suited for city life. It is in regular need of space and exercise and also requires regular brushing. They are not dogs for the casual hunter. Most are owned and used by foresters and game wardens.

History:
    The Bavarian Mountain Dog specialises in tracking injured big game following the traces of blood the prey loses after being shot. This hunting discipline comes from the Middle Ages. The weapons used then were imprecise and the animals were frequently wounded and not killed so the dogs were necessary to track them. Gaston Phébus wrote in 1387:
“It is a great amusement and a really nice hunt when you have a tracking dog and a good dog for the blood…”

The Germans were really meticulous in order not to lose any prey and developed a technique for which they bred resistant dogs with a great sense of smell, a strong bone structure, dropping ears and a steady temperament. These dogs were medium-sized and reliable. The Bavarian Mountain Dog was developed in the 19th century by crossbreeding specimens of the Hannoversche Schweißhund breed and hunting dogs from the Alps. The result was a hunting dog ideal for the work in the mountains. In 1912, the "Klub für Bayrische Gebirgsschweißhunde", (Club for Bavarian Mountain hound), was founded in Munich. Afterwards, this breed started gaining popularity in Austria and Hungary.

Bat


    Bats are mammals of the order Chiroptera (/kaɪˈrɒptərə/; from the Ancient Greek: χείρ - cheir, "hand" and Ancient Greek: πτερόν - pteron, "wing"whose forelimbs form webbed wings, making them the only mammals naturally capable of true and sustained flight. By contrast, other mammals said to fly, such as flying squirrels, gliding possums, and colugos, can only glide for short distances. Bats do not flap their entire forelimbs, as birds do, but instead flap their spread-out digits, which are very long and covered with a thin membrane or patagium.

    Bats are the second largest order of mammals (after the rodents), representing about 20% of all classified mammal species worldwide, with about 1,240 bat species divided into two suborders: the less specialized and largely fruit-eating megabats, or flying foxes, and the highly specialized and echolocating microbats. About 70% of bat species are insectivores. Most of the rest are frugivores, or fruit eaters. A few species, such as the fish-eating bat, feed from animals other than insects, with the vampire bats being hematophagous, or feeding on blood.

Bats are present throughout most of the world, with the exception of extremely cold regions. They perform the vital ecological roles of pollinating flowers and dispersing fruit seeds; many tropical plant species depend entirely on bats for the distribution of their seeds. Bats are economically important, as they consume insect pests, reducing the need for pesticides. The smallest bat is the Kitti's hog-nosed bat, measuring 29–34 mm (1.14–1.34 in) in length, 15 cm (5.91 in) across the wings and 2–2.6 g (0.07–0.09 oz) in mass. It is also arguably the smallest extant species of mammal, with the Etruscan shrew being the other contender. The largest species of bat are a few species of Pteropus (fruit bats or flying foxes) and the giant golden-crowned flying fox with a weight up to 1.6 kg (4 lb) and wingspan up to 1.7 m (5 ft 7 in).

The Mexican free-tailed bat is the fastest flying animal in horizontal flight.

Classification and evolution:
Bats are placental mammals. Bats were formerly thought to have been most closely related to the flying lemurs, treeshrews, and primates,[12] but recent molecular cladistics research indicates that they actually belong to Laurasiatheria, a diverse group also containing Carnivora and Artiodactyla.[13][14]

The two traditionally recognized suborders of bats are:

Megachiroptera (megabats)
Microchiroptera (microbats/echolocating bats)
Not all megabats are larger than microbats. The major distinctions between the two suborders are:

Microbats use echolocation; with the exception of the genus Rousettus, megabats do not.[15]
Microbats lack the claw at the second finger of the forelimb.[16][17]
The ears of microbats do not close to form a ring; the edges are separated from each other at the base of the ear.[17]
Microbats lack underfur; they are either naked or have guard hairs.[citation needed]
Megabats eat fruit, nectar, or pollen. Most microbats eat insects; others may feed on fruit, nectar, pollen, fish, frogs, small mammals, or the blood of animals. Megabats have well-developed visual cortices and show good visual acuity, while microbats rely on echolocation for navigation and finding prey.

The phylogenetic relationships of the different groups of bats have been the subject of much debate. The traditional subdivision between Megachiroptera and Microchiroptera reflects the view that these groups of bats have evolved independently of each other for a long time, from a common ancestor already capable of flight. This hypothesis recognized differences between microbats and megabats and acknowledged that flight has only evolved once in mammals. Most molecular biological evidence supports the view that bats form a single or monophyletic group.[18]

Researchers have proposed alternative views of chiropteran phylogeny and classification, but more research is needed.

In the 1980s, a hypothesis based on morphological evidence was offered that stated the Megachiroptera evolved flight separately from the Microchiroptera. The so-called flying primate hypothesis proposes that, when adaptations to flight are removed, the Megachiroptera are allied to primates by anatomical features not shared with Microchiroptera. One example is that the brains of megabats show a number of advanced characteristics that link them to primates. Although recent genetic studies strongly support the monophyly of bats,[19] debate continues as to the meaning of available genetic and morphological evidence.[20]

Genetic evidence indicates that megabats originated during the early Eocene and should be placed within the four major lines of microbats.[21]

Consequently, two new suborders based on molecular data have been proposed. The new suborder of Yinpterochiroptera includes the Pteropodidae, or megabat family, as well as the families Rhinolophidae, Hipposideridae, Craseonycteridae, Megadermatidae, and Rhinopomatidae[22] The other new suborder, Yangochiroptera, includes all of the remaining families of bats (all of which use laryngeal echolocation). These two new suborders are strongly supported by statistical tests. Teeling (2005) found 100% bootstrap support in all maximum likelihood analyses for the division of Chiroptera into these two modified suborders. This conclusion is further supported by a 15-base-pair deletion in BRCA1 and a seven-base-pair deletion in PLCB4 present in all Yangochiroptera and absent in all Yinpterochiroptera.[22] Perhaps most convincingly, a phylogenomic study by Tsagkogeorga et al (2013) showed that the two new proposed suborders were supported by analyses of thousands of genes.[21]

The chiropteran phylogeny based on molecular evidence is controversial because microbat paraphyly implies that one of two seemingly unlikely hypotheses occurred. The first suggests that laryngeal echolocation evolved twice in Chiroptera, once in Yangochiroptera and once in the rhinolophoids.[23][24] The second proposes that laryngeal echolocation had a single origin in Chiroptera, was subsequently lost in the family Pteropodidae (all megabats), and later evolved as a system of tongue-clicking in the genus Rousettus.[25]


Common pipistrelle, Pipistrellus pipistrellus
Analyses of the sequence of the "vocalization" gene, FoxP2, were inconclusive as to whether laryngeal echolocation was secondarily lost in the pteropodids or independently gained in the echolocating lineages.[26] However, analyses of the "hearing" gene, Prestin, seemed to favor the independent gain in echolocating species rather than a secondary loss in the pteropodids.[27]

In addition to Yinpterochiroptera and Yangochiroptera, the names Pteropodiformes and Vespertilioniformes have also been proposed for these suborders.[28][29] Under this new proposed nomenclature, the suborder Pteropodiformes includes all extant bat families more closely related to the genus Pteropus than the genus Vespertilio, while the suborder Vespertilioniformes includes all extant bat families more closely related to the genus Vespertilio than to the genus Pteropus.

Little fossil evidence is available to help map the evolution of bats, since their small, delicate skeletons do not fossilize very well. However, a Late Cretaceous tooth from South America resembles that of an early microchiropteran bat. Most of the oldest known, definitely identified bat fossils were already very similar to modern microbats. These fossils, Icaronycteris, Archaeonycteris, Palaeochiropteryx and Hassianycteris, are from the early Eocene period, 52.5 million years ago.[18] Archaeopteropus, formerly classified as the earliest known megachiropteran, is now classified as a microchiropteran.

Bats were formerly grouped in the superorder Archonta, along with the treeshrews (Scandentia), colugos (Dermoptera), and the primates, because of the apparent similarities between Megachiroptera and such mammals. Genetic studies have now placed bats in the superorder Laurasiatheria, along with carnivorans, pangolins, odd-toed ungulates, even-toed ungulates, and cetaceans.[30] A recent study by Zhang et al. places Chiroptera as a sister taxon to the clade Perissodactyla (which includes horses and other odd-toed ungulates).[31] However, the first phylogenomic analysis of bats shows that they are not sisters to Perissodactyla, instead they are sisters to a larger group that includes ungulates and carnivores.[21]